Brusatte, S. L., Averianov, A., Sues, H.-D., Muir, A. & Butler, I. B. New tyrannosaur from the mid-Cretaceous of Uzbekistan clarifies evolution of big physique sizes and superior senses in tyrant dinosaurs. Proc. Natl Acad. Sci. USA 113, 3447–3452 (2016).
Nesbitt, S. J. et al. A mid-Cretaceous tyrannosauroid and the origin of North American end-Cretaceous dinosaur assemblages. Nat. Ecol. Evol. 3, 892–899 (2019).
Zanno, L. E. et al. Diminutive fleet-footed tyrannosauroid narrows the 70-million-year hole within the North American fossil report. Commun. Biol. 2, 64 (2019).
Lengthy, J. A. & McNamara, Okay. J. in Evolutionary Change and Heterochrony (ed. McNamara, Okay. J.) 151–168 (Wiley, 1995).
Lengthy, J. A. & McNamara, Okay. J. in Dinofest Worldwide. Proceedings of a Symposium held at Arizona State College (eds. Wolberg, D. L. et al.) 113–123 (Academy of Pure Sciences, 1997).
Erickson, G. M. Assessing dinosaur development patterns: a microscopic revolution. Traits Ecol. Evol. 20, 677–684 (2005).
Erickson, G. M. et al. Gigantism and comparative life-history parameters of tyrannosaurid dinosaurs. Nature 430, 772–775 (2004).
Cullen, T. M. et al. Osteohistological analyses reveal numerous methods of theropod dinosaur body-size evolution. Proc. R. Soc. B 287, 20202258 (2020).
Voris, J. T. et al. Two exceptionally preserved juvenile specimens of Gorgosaurus libratus (Tyrannosauridae, Albertosaurinae) present new perception into the timing of ontogenetic adjustments in tyrannosaurids. J. Vertebr. Paleontol. 41, e2041651 (2022).
D’Emic, M. D. et al. Developmental methods underlying gigantism and miniaturization in non-avialan theropod dinosaurs. Science 379, 811–814 (2023).
Holtz Jr, T. R. in Mesozoic Vertebrate Life (eds Tanke, D. H. et al.) 64–83 (Indiana Univ. Press, 2001).
Brusatte, S. L., Carr, T. D., Erickson, G. M., Bever, G. S. & Norell, M. A. A protracted-snouted, multihorned tyrannosaurid from the Late Cretaceous of Mongolia. Proc. Natl Acad. Sci. USA 106, 17261–17266 (2009).
Brusatte, S. L. et al. Tyrannosaur paleobiology: new analysis on historic exemplar organisms. Science 329, 1481–1485 (2010).
Loewen, M. A., Irmis, R. B., Sertich, J. J. W., Currie, P. J. & Sampson, S. D. Tyrant dinosaur evolution tracks the rise and fall of Late Cretaceous oceans. PLoS ONE 8, e79420 (2013).
Lü, J. et al. A brand new clade of Asian Late Cretaceous long-snouted tyrannosaurids. Nat. Commun. 5, 3788 (2014).
Brusatte, S. L. & Carr, T. D. The phylogeny and evolutionary historical past of tyrannosauroid dinosaurs. Sci. Rep. 6, 20252 (2016).
Carr, T. D., Varricchio, D. J., Sedlmayr, J. C., Roberts, E. M. & Moore, J. R. A brand new tyrannosaur with proof for anagenesis and crocodile-like facial sensory system. Sci. Rep. 7, 44942 (2017).
Voris, J. T., Therrien, F., Zelenitsky, D. Okay. & Brown, C. M. A brand new tyrannosaurine (Theropoda:Tyrannosauridae) from the Campanian Foremost Formation of Alberta, Canada, offers perception into the evolution and biogeography of tyrannosaurids. Cretac. Res. 110, 104388 (2020).
Dalman, S. G. et al. An enormous tyrannosaur from the Campanian–Maastrichtian of southern North America and the evolution of tyrannosaurid gigantism. Sci. Rep. 14, 22124 (2024).
Delcourt, R. & Grillo, O. N. Tyrannosauroids from the Southern Hemisphere: implications for biogeography, evolution, and taxonomy. Palaeogeogr. Palaeoclimatol. Palaeoecol. 511, 379–387 (2018).
Gignac, P. M. & Erickson, G. M. The biomechanics behind excessive osteophagy in Tyrannosaurus rex. Sci. Rep. 7, 2012 (2017).
Tanaka, Okay., Anvarov, O. U. O., Zelenitsky, D. Okay., Ahmedshaev, A. S. & Kobayashi, Y. A brand new carcharodontosaurian theropod dinosaur occupies apex predator area of interest within the early Late Cretaceous of Uzbekistan. R. Soc. Open Sci. 8, 210923 (2021).
Perle, A. On the invention of Alectrosaurus from the Late Cretaceous of Mongolia. Probl. Mong. Geol. 3, 104–113 (1977).
Carr, T. D. A reappraisal of tyrannosauroid fossils from the Iren Dabasu Formation (Coniacian–Campanian), Internal Mongolia, Individuals’s Republic of China. J. Vertebr. Paleontol. 42, e2199817 (2023).
Mader, B. J. & Bradley, R. L. A redescription and revised analysis of the syntypes of the Mongolian tyrannosaur Alectrosaurus olseni. J. Vertebr. Paleontol. 9, 41–55 (1989).
Gilmore, C. W. On the dinosaurian fauna of the Iren Dabasu Formation. Bull. Am. Mus. Nat. Hist. 67, 23–95 (1933).
Van Itterbeeck, J., Horne, D. J., Bultynck, P. & Vandenberghe, N. Stratigraphy and palaeoenvironment of the dinosaur-bearing Higher Cretaceous Iren Dabasu Formation, Internal Mongolia, Individuals’s Republic of China. Cretac. Res. 26, 699–725 (2005).
Holtz Jr, T. R. in The Dinosauria (eds Weishampel, D. B. et al.) 111–136 (Univ. California Press, 2004).
Brochu, C. A. Osteology of Tyrannosaurus rex: insights from an almost full skeleton and high-resolution computed tomographic evaluation of the cranium. J. Vertebr. Paleontol. 22, 1–138 (2003).
Averianov, A. & Sues, H.-D. Skeletal stays of Tyrannosauroidea (Dinosauria: Theropoda) from the Bissekty Formation (Higher Cretaceous: Turonian) of Uzbekistan. Cretac. Res. 34, 284–297 (2012).
von Huene, F. Das natürliche system der Saurischia. Zentralbl. Mineral. Geol. Paläontol. B 1914, 154–158 (1914).
Gauthier, J. A. Saurischian monophyly and the origin of birds. Mem. Calif. Acad. Sci. 8, 1–55 (1986).
Osborn, H. F. Tyrannosaurus and different Cretaceous carnivorous dinosaurs. Bull. Am. Mus. Nat. Hist. 21, 259–265 (1905).
Tsuihiji, T., Watabe, M., Tsogtbaatar, Okay., Barsbold, R. & Suzuki, S. A tyrannosauroid frontal from the Higher Cretaceous (Cenomanian-Santonian) of the Gobi Desert, Mongolia. Vertebr. Palasiat. 50, 102–110 (2012).
Averianov, A. & Sues, H.-D. Correlation of Late Cretaceous continental vertebrate assemblages in center and central Asia. J. Stratigr. 36, 462–485 (2012).
Brusatte, S. L., Carr, T. D. & Norell, M. A. The osteology of Alioramus, a gracile and long-snouted tyrannosaurid (Dinosauria: Theropoda) from the Late Cretaceous of Mongolia. Bull. Am. Mus. Nat. Hist. 366, 1–197 (2012).
Foster, W. et al. The cranial anatomy of the long-snouted tyrannosaurid dinosaur Qianzhousaurus sinensis from the Higher Cretaceous of China. J. Vertebr. Paleontol. 41, e1999251 (2022).
Carr, T. D. A high-resolution development sequence of Tyrannosaurus rex obtained from a number of strains of proof. PeerJ 8, e9192 (2020).
Carr, T. D. Craniofacial ontogeny in Tyrannosauridae (Dinosauria, Coelurosauria). J. Vertebr. Paleontol. 19, 497–520 (1999).
Currie, P. J. Allometric development in tyrannosaurids (Dinosauria: Theropoda) from the Higher Cretaceous of North America and Asia. Can. J. Earth Sci. 40, 651–665 (2003).
Currie, P. J. Cranial anatomy of tyrannosaurid dinosaurs from the Late Cretaceous of Alberta,Canada. Acta Palaeontol. Pol. 48, 191–226 (2003).
Tsuihiji, T. et al. Cranial osteology of a juvenile specimen of Tarbosaurus bataar (Theropoda, Tyrannosauridae) from the Nemegt Formation (Higher Cretaceous) of Bugin Tsav, Mongolia. J. Vertebr. Paleontol. 31, 497–517 (2011).
Xu, X. et al. Basal tyrannosauroids from China and proof for protofeathers in tyrannosauroids. Nature 431, 680–684 (2004).
Hutt, S., Naish, D., Martill, D. M., Barker, M. J. & Newbery, P. A preliminary account of a brand new tyrannosauroid theropod from the Wessex Formation (Early Cretaceous) of southern England. Cretac. Res. 22, 227–242 (2001).
Naish, D. & Cau, A. The osteology and affinities of Eotyrannus lengi, a tyrannosauroid theropod from the Wealden Supergroup of southern England. PeerJ 10, e12727 (2022).
Li, D., Norell, M. A., Gao, Okay.-Q., Smith, N. D. & Makovicky, P. J. A longirostrine tyrannosauroid from the Early Cretaceous of China. Proc. R. Soc. B 277, 183–190 (2010).
Voris, J. T., Zelenitsky, D. Okay., Therrien, F. & Currie, P. J. Reassessment of a juvenile Daspletosaurus from the Late Cretaceous of Alberta, Canada with implications for the identification of immature tyrannosaurids. Sci. Rep. 9, 17801 (2019).
Benson, R. B. J. New data on Stokesosaurus, a tyrannosauroid (Dinosauria: Theropoda) from North America and the UK. J. Vertebr. Paleontol. 28, 732–750 (2008).
Brusatte, S. & Benson, R. The systematics of Late Jurassic tyrannosauroids (Dinosauria: Theropoda) from Europe and North America. Acta Palaeontol. Pol. 58, 47–54 (2012).
Carr, T. D., Williamson, T. E. & Schwimmer, D. R. A brand new genus and species of tyrannosauroid from the Late Cretaceous (Center Campanian) Demopolis Formation of Alabama. J. Vertebr. Paleontol. 25, 119–143 (2005).
Brusatte, S. L., Benson, R. B. J. & Norell, M. A. The anatomy of Dryptosaurus aquilunguis (Dinosauria: Theropoda) and a evaluate of Its tyrannosauroid affinities. Am. Mus. Novit. 3717, 1–53 (2011).
Currie, P. J., Hurum, R. H. & Sabath, Okay. Cranium construction and evolution in tyrannosaurid dinosaurs. Acta Palaeontol. Pol. 48, 227–234 (2003).
Carr, T. D. & Williamson, T. E. Variety of late Maastrichtian Tyrannosauridae (Dinosauria: Theropoda) from western North America. Zool. J. Linn. Soc. 142, 479–523 (2004).
Goloboff, P. A. & Morales, M. E. TNT model 1.6, with a graphical interface for MacOS and Linux, together with new routines in parallel. Cladistics 39, 144–153 (2023).
Groh, S. S., Upchurch, P., Barrett, P. M. & Day, J. J. Learn how to date a crocodile: estimation of neosuchian clade ages and a comparability of 4 time-scaling strategies. Palaeontology 65, e12589 (2022).
Bapst, D. W. A stochastic rate-calibrated methodology for time-scaling phylogenies of fossil taxa. Strategies Ecol. Evol. 4, 724–733 (2013).
Lloyd, G. T., Bapst, D. W., Friedman, M. & Davis, Okay. E. Probabilistic divergence time estimation with out department lengths: relationship the origins of dinosaurs, avian flight and crown birds. Biol. Lett. 12, 20160609 (2016).
Revell, L. J. phytools 2.0: an up to date R ecosystem for phylogenetic comparative strategies (and different issues). PeerJ 12, e16505 (2024).
Voris, J. T et al. Information for ‘A brand new Mongolian tyrannosauroid and the evolution of Eutyrannosauria’. Dryad https://doi.org/10.5061/dryad.fj6q5744h (2025).